Syncing client crashes with "Network Error:99"

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Files created on the client will not be synced to the server, client log show “Network Error 99”

Everything freshly setup on Friday in a new Ubuntu 22.04 LXD container, mostly following

Everything else seems to be working fine.

Apache proxy config on the gateway container is

<VirtualHost *:443>
  ServerName cloud.example.org
  ServerAdmin webmaster@example.org

  AllowEncodedSlashes On

  RewriteEngine On
  RewriteRule ^/\.well-known/carddav https://%{SERVER_NAME}/remote.php/dav [R=301,L]
  RewriteRule ^/\.well-known/caldav https://%{SERVER_NAME}/remote.php/dav [R=301,L]
  RewriteRule ^/\.well-known/webfinger https://%{SERVER_NAME}/index.php/.well-known/webfinger [R=301,L]
  RewriteRule ^/\.well-known/nodeinfo https://%{SERVER_NAME}/index.php/.well-known/nodeinfo [R=301,L]

  Include /etc/letsencrypt/options-ssl-apache.conf
  Header always set Strict-Transport-Security "max-age=15552000; includeSubDomains"

  SetEnvIf Authorization "(.*)" HTTP_AUTHORIZATION=$1
  ProxyPreserveHost On
  ProxyPass / http://10.90.60.236:80/
  ProxyPassReverse / http://10.90.60.236:80/
  SSLCertificateFile /etc/letsencrypt/live/example.org/fullchain.pem
  SSLCertificateKeyFile /etc/letsencrypt/live/example.org/privkey.pem
</VirtualHost>